McCullough, E. L., Buzatto, B. A., and Simmons, L. W. 2018. Population density mediates the interaction between pre- and postmating sexual selection. Evolution 72: 893-905. PDF

McCullough, E. L. and Emlen, D. J. 2018. The research bias is unfortunate but also unsurprising: a comment on Tinghitella et al. Behavioral Ecology in press. PDF

McCullough, E. L., Buzatto, B. A., and Simmons, L.W. 2017. Benefits of polyandry: molecular evidence from field-caught dung beetles. Molecular Ecology 26: 3546-3555. PDF

McCullough, E. L., Miller, C. W., and Emlen, D. J. 2016. Why sexually selected weapons are not ornaments. Trends in Ecology and Evolution 31: 742-751. (cover article) PDF

McCullough, E. L. and Simmons, L. W. 2016. Selection on male physical performance during male-male competition and female choice. Behavioral Ecology 27: 1288-1295. PDF

McCullough, E. L., Ledger, K. J., O’Brien, D. M., and Emlen, D. J. 2015. Variation in the allometry of exaggerated rhinoceros beetle horns. Animal Behaviour 109: 133-140. PDF

McCullough, E. L., Ledger, K. J., and Moore, T. Y. 2015. Variation in cross-sectional horn shape within and among rhinoceros beetle species. Biological Journal of the Linnean Society 115: 810-817. PDF

McCullough, E. L., Tobalske, B. W., and Emlen, D. J. 2014. Structural adaptations to diverse fighting styles in sexually selected weapons. Proceedings of the National Academy of Sciences 111: 14484-14488. PDF

McCullough, E. L. 2014. Mechanical limits to maximum weapon size in a giant rhinoceros beetle. Proceedings of the Royal Society B 281: 20140696. PDF

Johns, A., Gotoh, H., McCullough, E. L., Emlen, D. J., and Lavine, L. C. 2014. Heightened condition-dependent growth of sexually selected weapons in the rhinoceros beetle, Trypoxylus dichotomus (Coleoptera: Scarabaeidae). Integrative and Comparative Biology 54: 614-621. PDF

Najera, D. A., McCullough, E. L., Jander, R. 2014. Honeybees use celestial and/or terrestrial compass cues for inter-patch navigation. Ethology 120: 1-9.

McCullough, E. L. and Emlen, D. J. 2013. Evaluating the costs of a sexually selected weapon: big horns at a small price. Animal Behaviour 86: 977-985. PDF

McCullough, E. L. and Zinna, R. A. 2013. Sensilla density corresponds to the regions of the horn most frequently used during combat in the giant rhinoceros beetle Trypoxylus dichotomus (Coleoptera: Scarabaeidae: Dynastinae). Annals of the Entomological Society of America 106: 518-523. PDF

McCullough, E.L. and Tobalske, B.W. 2013. Elaborate horns in a giant rhinoceros beetle incur negligible aerodynamic costs. Proceedings of the Royal Society B 280: 20130197. PDF

McCullough, E. L. 2013. Using radio telemetry to assess movement patterns in a giant rhinoceros beetle: Are there differences among majors, minors, and females? Journal of Insect Behavior 26: 51-56. PDF

McCullough, E. L., Weingarden, P. R., Emlen, D. J. 2012. Costs of elaborate weapons in a rhinoceros beetle: How difficult is it to fly with a big horn? Behavioral Ecology 23: 1042-1048. PDF

Najera, D. A., McCullough, E. L., Jander, R. 2012. Interpatch foraging in honeybees: Rational decision making at secondary hubs based upon time and motivation. Animal Cognition 15: 1195- 1203.

Bai, M., McCullough, E., Song, K-Q., Liu, W-G., Yang, X-K. 2011. Evolutionary constraints in hind wing shape in Chinese dung beetles (Coleoptera: Scarabaeinae). PLoS ONE 6: e21600.

McCullough, E., Wright, K. M., Alvarez, A., Clark, C. P., Rickoll, W. L., and Madlung, A. 2010. Photoperiod-dependent floral reversion in the natural allopolyploid Arabidopsis suecica. New Phytologist 186: 239-250.